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Tariq Stévart, Ph.D.
Taking a picture of a new orchid species in Cameroon
Tariq Stévart, Ph.D.
Scientist and Coordinator of West
and Central Africa Program
Africa and Madagascar


Research Associate, Université Libre de Bruxelles, Belgium and at the Botanic Garden Meise

St. Louis address: Missouri Botanical Garden
4344 Shaw Blvd.
St. Louis, MO 63110

Belgium address:
Herbarium of the Université Libre de Bruxelles, Faculté des Sciences, Campus de la Plaine - CP 265
Boulevard du Triomphe, ACC.2, 1050 Bruxelles
Botanic Garden Meise, Domein van Bouchout, Nieuwelaan 38, B-1860, Meise, Belgium
Phone (Belgium): 32 494632253

Gabon address:
WCS Gabon, Derrière Le Camp De Gaulle, Rue L'Arche De L'Alliance Libreville, Gabon
Phone (Gabon): 241 07350019

Ph.D., Université Libre de Bruxelles, Belgium, 2003
M.S., Université Libre de Bruxelles, Belgium, 2000
B.S., Université Libre de Bruxelles, Belgium, 1998
Taxonomy and Phylogeny of African angraecoid Orchids Flora and vegetation of West Central Africa Biogeography and diversity of the African forests Application of floristic and vegetation data to conservation planning and management Conservation of the African flora and ex situ conservation of African orchids IUCN Red Listing of African and Malagasy flora

Web Pages
The Central Africa orchid shadehouse network
For Dr. Stévart's complete CV with list of publications, click here (Adobe Acrobat Reader required).
Research Emphases Taxonomy and biogeography of Orchidaceae and Rubiaceae of Africa with Bonaventure Sonké (Université de Yaoundé I, Cameroon) and Vincent Droissart (IRD, France). Description of new taxa (with D. Geerinck, P.J. Cribb, V. Droissart & B. Sonké); revisions of selected genera: Chamaengis (with V. Droissart & B. Sonké), Angraecum (with V. Droissart), Ancistrorhynchus (with D. Geerinck). Phylogenetic study of the angraecoid orchids (with Gregory M. Plunkett, NYBG and Claire Micheneau ULB). Ex situ and in situ conservation of orchids (with V. Droissart, B. Sonké, M. Simo) and endemic African plants (with Gretchen Walters, IUCN and Steven Dessein, BR). Diversity of the African forests with Olivier Hardy (ULB, Belgium). Development of the IUCN Red Listing in Central Africa with Steven Dessein (BR), David KenFack (SI) and Roy Gereau (MBG). Flora of Gabon: inventory in the Massif du Chaillu, Waka National Park and Monts de Cristal; identification of new protected areas (with Gretchen Walters (IUCN). Sao Tome and Principe: Catalogue of the endemic Vascular Plants (with G. Joffroy, E. Figeiredo & F. do Oliveira). West Central Africa: biogeography of inselbergs (with I. Parmentier); ecology of epiphytes on inselbergs in relation with surrounding forests.
Selected Publications Stévart T., Dauby G., Lowry PP., Blach-Overgaard A., Droissart V., Harris DJ., Mackinder BA., Schatz GE., Sonké B., Sosef MSM., Svenning J.-C., Wieringa J. J., Couvreur T. L. P. 2019. A third of the tropical African flora is potentially threatened with extinction, Science Advances, 5,11,eaax9444, American Association for the Advancement of Science. Dagallier L-P., Janssens S.B., Dauby G., Blach-Overgaard A., Mackinder B.A., Droissart V., Svenning, J-C., Sosef M. SM, Stévart T., Harris D. J., 2019. Cradles and museums of generic plant diversity across tropical Africa, New Phytologist. Wiley Online Library. Sosef M. SM, Bidault E., Boupoya A., Lachenaud O., Stévart T. 2019. Novitates Gabonenses 91: additions to the grass flora of Gabon, Plant Ecology and Evolution, 152, 1: 99-105. Lissambou B.-J., Couvreur T. LP, Atteke C., Stévart T., Piñeiro R., Dauby G., Sonké B., M’batchi B., Hardy O.J. 2019. Species delimitation in the genus Greenwayodendron based on morphological and genetic markers reveals new species, Taxon, 68: 442-454. Droissart V., Lachenaud O., Dauby G., Dessein S., Kamdem G., Nguembou K., Simo-Droissart M., Stévart T., Taedoumg H., Sonké B. 2019. Mine versus Wild: a plant conservation checklist of the rich Iron-Ore Ngovayang Massif Area (South Cameroon). Plant Ecology and Evolution,152,1, 8-29. Simo-Droissart M., Stévart T. 2019. A new combination in Dolabrifolia (Angraecinae, Orchidaceae). Phytotaxa, 404, 7, 295-296. D haijère T. , Mardulyn P., Ling Dong, W., Plunkett G.M., Mardulyn P., Droissart, V., Simo-Droissart, M., Stévart T.. 2019. Molecular phylogeny and taxonomic synopsis of the angraecoid genus Ypsilopus (Orchidaceae, Vandeae). Taxon, 68: 455-470. Lissambou B.-J., Hardy O.J. Atteke C., Stévart T., Dauby G., M’batchi B., Sonké B., Couvreur T. LP. 2018. Taxonomic revision of the African genus Greenwayodendron (Annonaceae). PhytoKeys, 114, 55-93. Descourvières P., Farminhão J., Droissart V., Dubuisson J.Y., Simo-Droissart, M., Stévart T. 2018. A new genus of angraecoid orchids (Orchidaceae: Angraecinae) with highly distinctive pollinaria morphology, including three new species from tropical West and Central Africa. Phytotaxa 373 (2): 099–120. Ikabanga D.U., Koffi K.G., O.M., M'batchi B., Hardy O. J., Stévart T. 2019. Révision taxonomique du genre Santiria (Burseraceae) en Afrique tropicale. Candollea, 74 (2): 115-130. Pauwels O.S.G., Bahaa-el-din L., Albret J.-L., Carlino P., Giannuzzi F., Chirio L., Gillet J.-F., Poirier E., Stévart T. 2018. Miscellanea Herpetologica Gabonica XIV. Bulletin of the Chicago Herpetological Society 53(9), 185-190. Lachenaud O., Stévart T., Boupoya A., Texier N., Dauby G., Bidault E. 2018. Novitates Gabonenses 88: additions to the flora of Gabon and new records of little-known species. Plant ecology and evolution, 151, 3, 393-422. Texier N., Deblauwe V., Stévart T., Sonké B., Simo-Droissart M., Djuikouo M.-N., Zemagho L., Guiakam C., Azandi L., Kamdem G., Kamdem N., Mayogo S., Droissart V. 2018. Spatio-temporal patterns orchids flowering in Cameroonian rainforests. International Journal of Biometeorology. Simo-Droissart, M., Plunkett, G. M., Droissart, V., Edwards, M. B., Farminhão, J. N. M., Jecmenica, V., D haijère, T., Lowry II, P.P., Sonké, B., Micheneau, C., Carlsward, B. S. , Azandi, L., Verlyndes, S., Hardy, O. J., Martos, F., Bytebier, B., Fischer, E., Stévart, T. 2018. New phylogenetic insights toward developing a natural generic classification of African angraecoid orchids (Vandeae, Orchidaceae). Molecular Phylogenetics and Evolution, 126 : 241-249. Simo-Droissart, M., Stévart, T., Sonké, B., Mayogo, S., Kamdem, N. G., Droissart, V. 2018. New taxonomic and conservation status of Ossiculum (Vandeae, Orchidaceae), a highly threatened and narrow-endemic angraecoid orchid from Central Africa. PhytoKeys , 98 : 85-97. Simo-Droissart, M., Sonké, B., Droissart, V., Stévart, T. 2018. Afropectinariella (Vandeae, Orchidaceae), a new genus of the Angraecum alliance. PhytoKeys , 96 : 79-86. Droissart, V., Dauby, G., Hardy, O. J., Deblauwe, V., Harris, D. J., Janssens, S., Mackinder, B., Blach-Overgaard, A., Sonké, B., Sosef, M. S. M., Stévart, T., Svenning, J.-C., Wieringa, J. J., Couvreur, T. P. L. 2018. Beyond trees: Biogeographical regionalization of tropical Africa. Journal of Biogeography, 45 (5) : 1153-1167. Farminhão, J. N. M., Meerts, P., Descourvières, P., Droissart, V., Simo-Droissart, M., Stévart, T. 2018. A revised concept of Rhipidoglossum (Angraecinae, Orchidaceae). Phytotaxa , 349 (3) : 247-256. Tania. Verlyndes, S., D Haese, C. A., Plunkett, G. M., Simo-Droissart, M., Edwards, M., Droissart, V., Stévart, T. 2018. Molecular phylogeny of the genus Bolusiella (Orchidaceae, Angraecinae). Plant Systematics and Evolution, 304 (2) : 269-279. Slik F. et al. 2017. A Phylogenetic Classification Of The World’s Tropical Forests. Proceedings of the National Academy of Sciences 115(8). Lachenaud O., Schatz G.E., Dauby G., Stévart T. (2017). Two new species of Diospyros (Ebenaceae) from Central Africa Plant Ecology and Evolution. 150 (2): 217–224. Dauby, G., Stévart, T., Droissart, V., Cosiaux, A., Deblauwe, V., Simo-Droissart, M., Sosef, M. S. M., Lowry II, P.P., Schatz, G. E., Gereau, R. E., Couvreur, T. P. L. 2017. ConR: An R package to assist large-scale multispecies preliminary conservation assessments using distribution data. Ecology and Evolution, 7 (24) : 11292-11303. Jecmenica V., Droissart V., Akouangou E., Nyangala C., Bakita B., Biteau J. P., Stévart T. (2017). Taxonomy of Atlantic Central African orchids, 6: three new species of Angraecum sect. Afrangraecum (Orchidaceae, Angraecinae) from Gabon and Sao Tomé. Phytotaxa, 323 (2), 143-158. ISSN 1179-3155. Ikabanga D., Stévart T., Koffi, Kouamé G., Monthe Kameni, F., Nzigoudoubindou, E.,; Dauby G.; Souza A.; M’Batchi B., Hardy O.J. 2017. Combining morphology and population genetic analysis uncover species delimitation in the widespread African tree genus Santiria (Burseraceae). Phytotaxa, 321, 2: 166-180. Sosef M., Dauby G., Blach-Overgaard A., van der Burgt X., Catarino L., Damen T., Deblauwe V., Dessein S., Dransfield J., Droissart V. , Duarte M.C., Engledow H., Fadeur G., Figueira R., Gereau R., Hardy O.J., Harris D.J., de Heij J., Janssens S., Klomberg Y., Ley A.C., Mackinder B.A., Meerts P., van de Poel J.L., Sonké B., Stévart T., Stoffelen P., Svenning J.C., Sepulchre P., Zaiss R., Wieringa J.J., Couvreur T.L.P. 2017. Exploring the Floristic Diversity of Tropical Africa BMC Biol 15 (1): 15. Vissers J, Bosch FV, Bogaerts A, Cocquyt C, Degreef J, Diagre D, de Haan M, De Smedt S, Engledow H, Ertz D, Fabri R, Godefroid S, Hanquart N, Mergen P, Ronse A, Sosef M, Stévart T., Stoffelen P, Vanderhoeven S, Groom Q. 2017. Scientific user requirements for a herbarium data portal. PhytoKeys Dauby, G., Zaiss, R., Blach-Overgaard, A., Catarino, L., Damen, T., Deblauwe, V., Dessein, S., Dransfield, J., Droissart, V., Duarte, M. C., Engledow, H., Fadeur, G., Figueira, R., Gereau, R. E., Hardy, O. J., Harris, D. J., de Heij, J., Janssens, S., Klomberg, Y., Ley, A. C., Mackinder, B., Meerts, P., van de Poel, J. L., Sonké, B., Sosef, M. S. M., Stévart, T., Stoffelen, P., Svenning, J.-C., Sepulchre, P., van der Burgt, X., Couvreur, T. P. L. 2016. RAINBIO: a mega-database of tropical African vascular plants distributions. PhytoKeys, 74 (1). Deblauwe V., Droissart V., Bose R., Sonké B., Blach-Overgaard A., Svenning J.-C., Wieringa J. J., Ramesh B. R., Stévart T. & Couvreur T. L. P. 2016. Remotely sensed temperature and precipitation data improve species distribution modelling in the tropics. Global Ecology and Biogeography, (Global Ecol. Biogeogr.). 25 (4) 443–454. Bidault E., Lowry II P.P., Stévart T. 2016. Polystachya orophila (Orchidaceae, Polystachynae), a new species from tropical West Africa, and clarification on the nomenclature and taxonomy of P. microbambusa. Phytotaxa. 260(3): 247–257. Azandi L., Stévart T., Sonké B., Simo-Droissart M., Avana L., Droissart V. 2016. Synoptic revision of the genus Cyrtorchis Schltr. (Angraecinae, Orchidaceae) in Central Africa, with a new species endemic to submontane vegetation. Phytotaxa. 34: 1–64. Verlynde S., Ramandimbisoa B., Bosser J. & Stévart T. 2016. Contribution à l’étude des Orchidaceae de Madagascar. XXXVII. Deux Pectinariella (Orchidaceae) nouveaux de Madagascar et redélimitation de Angraecum pterophyllum H. Perrier. Adansonia, 38(2), 219-232. Simo-Droissart M., Sonké B., Droissart V., Lowry II P.P., Micheneau C., Hardy O.J., Plunkett G.M., Stévart T. 2016. Morphometrics and molecular phylogenetics of Angraecum section Dolabrifolia (Orchidaceae, Angraecinae) 280(2) : 81–115. Ječmenica V., Droissart V., Noret N., Stévart T. 2016. Taxonomy of Atlantic Central African orchids 5. A new species of Angraecum sect. Conchoglossum (Orchidaceae, Angraecinae) from Gabon and Cameroon. PhytoKeys 61: 61-71. Slik F. et al. 2015. An estimate of the number of tropical tree species. PNAS, 12(24):7472–7477, Bastin, J.-F. et al. 2015. Seeing Central African forests through their largest trees. Sci. Rep. 5, 13156; D'haijère T., Leguil S., Meerts P., Stévart T. 2015. Taxonomy of Atlantic Central African orchids 4. A new species of Tridactyle (Vandeae, Epidendroideae, Orchidaceae) from Gabon and Equatorial Guinea. Phytotaxa, Phytotaxa, 212 (2): 141–148. Walters G., Ngagnia Ndjabounda E., Ikabanga D., Biteau J. P., Hymas O., White L. J. T., Ndong Obiang A.-M., Ndong Ondo P., Jeffery K. J., Lachenaud O. and T. Stévart. 2015. Peri-urban conservation in the Mondah forest of Libreville, Gabon: Red List assessments of endemic plant species, and avoiding protected area downsizing. Oryx, available on CJO2015. Maréchal C. et al. 2014. Les forêts du bassin du Congo – État des Forêts 2013, Chapter: Conservation et gestion de la biodiversité, Publisher: Weyrich Édition, Editors: de Wasseige C., Flynn J., Louppe D., Hiol Hiol F, Mayaux P, pp. 67–96. Stévart T., Biteau J.P., Cawoy V., Droissart V. 2014. Taxonomy of Atlantic Central African orchids 3. A new species of Bulbophyllum Thouars (Orchidaceae) from the Monts de Cristal, Gabon. Phytotaxa 177 (1): 026–034 Dauby G., Duminil J., Heuertz M., Koffi G.K., Stévart T. & Hardy O.J. 2014. Congruent phylogeographical patterns of eight tree species in Atlantic Central Africa provide insights into the past dynamics of forest cover, Molecular Ecology, 23: 2299–2312. Dauby G., Hardy O.J., Leal M., Breteler F.J. & Stévart T. 2014. Drivers of tree diversity in tropical rain forests: new insights from a comparison between littoral and hilly landscapes of Central Africa, Journal of Biogeography, 41(3) 574-586. Simo-Droissart M., Sonké B., Droissart V., Micheneau C., Lowry II P.P., Plunkett G.M., Hardy O.J., Stévart T. 2014. Taxonomic revision of Continental African species of the section Pectinaria (Angraecum, Orchidaceae). Systematic Botany 39(3):725-739. Droissart V, Cribb P, Simo-Droissart M, Stévart T. 2014. Taxonomy of Atlantic Central African orchids 2. A second species of the rare genus Distylodon (Orchidaceae, Angraecinae) collected in Cameroon. PhytoKeys 36: 27-34. Descourvières P., Dubuisson J.Y., Droissart V., Cribb P.J., Cawoy V., Simo-Droissart M., Sonké B., Stévart T. 2013 A new species of Rhipidoglossum (Orchidaceae) from Equatorial Guinea and Cameroon. Plant Ecology and Evolution 146 (3):389-394. Lachenaud O., Droissart V., Dessein S., Stévart T., Simo M., Lemaire B., Taedoumg H., Sonké B. 2013. New records for the flora of Cameroon, including a new species of Psychotria (Rubiaceae) and range extensions for some rare species. Plant Ecology and Evolution 146 (1):121-133. Lachenaud O., Stévart T., Ikabanga D., Ngagnia Ndjabounda E. & Gretchen Walters. 2013. Les forêts littorales de la région de Libreville (Gabon) et leur importance pour la conservation: description d’un nouveau Psychotria (Rubiaceae) endémique Plant Ecology and Evolution 146 (1): 68–74. Simo-Droissart M., Micheneau C., Sonké B., Droissart V., Plunkett G.M., lowry II P.P., Hardy O.J., Stévart T. 2013. Morphometrics and molecular phylogenetics of the continental African species of Angraecum section Pectinaria (Orchidaceae). Plant Ecology and Evolution 146 (3):295-309. Verlynde S., Dubuisson J.Y., Stévart T., Simo-Droissart M., Geerinck D., Sonké B., Cawoy V., Descourvières P., Droissart V. 2013.Taxonomic revision of the genus Bolusiella (Orchidaceae, Angraecinae) with a new species from Cameroon, Burundi and Rwanda. Phytotaxa 114 (1):1-22. Slik JWF, Paoli G, McGuire K,…, Stévart T., et al. (with 58 co-auteurs). 2013. Large trees drive forest aboveground biomass variation in moist lowland forests across the tropics. Global Ecology and Biogeography, 22: 1261-1271. Walters G., Parmentier I., Stévart T. 2012. Diversity and conservation value of Gabon's savanna and inselberg open vegetation: an initial gap analysis. Plant Ecology and Evolution 145: 46–54. Droissart V., Sonké B., Lowry II P. P., Pollard B. J. & Stévart T. 2012. Notes taxonomiques et chorologiques sur quelques Orchidaceae endémiques d’Afrique centrale atlantique. Adansonia, sér. 3, 34 (1): 23-35. Walters G., Dauby G., Stévart T., Dessein S., Niangadouma R., Lachenaud O. 2011. Novitates Gabonenses 80. Additions and corrections to the flora of Gabon. Edinburgh Journal of Botany 68 (3): 423–442 (2011). Droissart V., Hardy O.J., Sonké B., Dahdouh-Guebas F. & Stévart T. 2011. Heterogeneous Sampling to Assess Geographic Diversity Gradients? A Case-Study with Orchidaceae and Rubiaceae in Cameroon. Biotropica, 44, 44-52. Droissart V., Sonké B., Hardy O.J., Simo M., Taedoumg H., Nguembou K.C., Stévart T. 2011. Do plant families with contrasting functional traits show similar patterns of endemism in Central Africa? Biodiversity and Conservation. 20: 1507-1531. Figueiredo E., Paiva J., T., Oliveira F. & Smith G.F. 2011. An annotated catalogue of the flowering plants of São Tomé et Príncipe. Bothalia 41: 41–82. Janssens S.B., Fischer E., Stévart T. 2010. New insights on the origin of epiphytic Impatiens (Balsaminaceae) in West Central Africa based on molecular phylogenetic analysis. Taxon. 59(5): 1508–1518. Droissart V., Lowry P.P., Bosser J., Roberts D.L. & Stévart T. 2010. Correcting confusion regarding the identity, circumscription and synonymy of Angraecum gracile Thouars (Orchidaceae). Taxon, 59(5): 1578-1580. Stévart T., Cawoy V., Damen T. & Droissart V. 2010. Taxonomy of West Central African orchid (1): A new Angraecum sect. Pectinaria (Benth.) Schltr. (Orchidaceae) from Gabon and Equatorial Guinea. Systematic Botany. 35(2): 252–256. Stévart T., Delepierre G., Lebel J.P. & Geerinck G. 2010. Les Orchidaceae du Parc National de Nyungwe (Rwanda). In X. van der Burgt, J. van der Maesen & J.-M. Onana (eds). Sytématique et Conservation des plantes africaines. Royal Botanic Gardens, Kew. Proceeding du XVIIIème congrès de l’AETFAT, Yaoundé, Cameroun, 91-100.Stévart T., Delepierre G., Lebel J.P. & Geerinck G. 2010. Les Orchidaceae du Parc National de Nyungwe (Rwanda). In X. van der Burgt, J. van der Maesen & J.-M. Onana (eds). Sytématique et Conservation des plantes africaines. Royal Botanic Gardens, Kew. Proceeding du XVIIIème congrès de l’AETFAT, Yaoundé, Cameroun, 91-100. Simo M., Droissart V., Sonké B. & Stévart T. 2009. The Orchid Flora of the Mbam Minkom Hills (Yaounde, Cameroon). Belgian Journal of Botany,142(2): 111-123. Nguembou K. C., Ewedje E.-E. B. K., Droissart V., Stévart T. & Sonké B. 2009. Une espèce nouvelle de Bertiera (sous-genre Bertierella, Rubiaceae) d’Afrique centrale atlantique. Adansonia 31 (2): 397-406. Droissart, V., Sonké, B., Djuikouo, M.N., Nguembou, C., Parmentier, I. & Stévart T. 2009. Synopsis of the genus Chamaeangis (Orchidaceae) with two new taxa. Systematic Botany, 34(2): 285-296. Droissart V., Simo M., Sonké B., Cawoy V. & Stévart T. 2009. Le genre Stolzia (Orchidaceae) en Afrique centrale avec deux nouveaux taxons. Adansonia, sér. 3, 31 (1): 25-40. Droissart V., Sonké B., Simo M. & Stévart T. 2009. New orchid records from West Central Africa. Edinburgh Journal of Botany 66(1): 115–132. Dauby G., Parmentier I. & Stévart T. 2008. A new species of Afrothismia (Burmanniaceae) from Gabon, Nordic Journal of Botany 25: 268-271. Dauby G., Leal M. & Stévart T. 2008. Vascular plant checklist of the Pongara National Park, Gabon, Syst. Geogr. Pl. 78: 155-216. Stévart T., Parmentier I. & Droissart V. 2007. Deux espèces nouvelles de Polystachya (Orchidaceae) de Guinée Equatoriale. Adansonia, 29(1): 31-38. Droissart V., Sonké B. & Stévart T. 2006. Les Orchidaceae endémiques d’Afrique centrale atlantique présentes au Cameroun. Syst. Geogr. Pl. 76: 3-84. Stévart T., Ngok Banak L. & Sosef M. 2006. Synthèse des inventaires réalisés sur les Orchidaceae dans le cadre du Projet d'Evaluation des Aires Protégées du Gabon. In S.A. Ghazanfar & H.J. Beentje (eds.). Taxonomy and ecology of African Plants: their conservation and sustainable use. Proceedings du XVIIème Congrès de l'Aetfat, Royal Botanic Gardens Kew: 651-665. Sonké B., Esono P., Nguembou C., & Stévart T. 2005. Une nouvelle espèce de Bertiera Aubl. (Rubiaceae) du sous-genre Bertierella découverte en Guinée Équatoriale et au Cameroun. Adansonia 27(2): 309-315. Parmentier I., Stévart T. & Hardy O.J. 2005. The inselberg flora of Atlantic Central Africa. I. Determinants of species assemblages. Journal of Biogeography 32 (4): 685-696. Stévart T. & Cribb P. 2004. Five new Tridactyle (Orchidaceae) from Central Africa. Kew Bull. 59: 195-205. Stévart T. & Nguema N. 2004. Trois espèces et quatre combinaisons nouvelles de Polystachya (Orchidaceae) du Cameroun, de Guinée Equatoriale et du Gabon. Adansonia 26(2): 217-233. Stévart T. & Cribb P. 2004. New species and records of Orchidaceae from São Tomé and Príncipe. Kew Bull. 59(2): 77-86. Stévart T., Geerinck D., Parmentier I. & Lejoly J. 2003. A new species of Polystachyasect. Polychaete(Orchidaceae) from central Africa. Syst. Geogr. Pl. 73: 281-285. Stévart T. & Geerinck D. 2001. Diversité des Orchidaceae à São Tomé et Príncipe. Proceedings 16th Congress of A.E.T.F.A.T., 2000, Meise, Belgium. Syst. Geogr. Pl. 71: 565-573. Stévart T., Geerinck D. & Lejoly J. 2000. Liste des Orchidaceae de São Tomé et Príncipe. Acta Bot. Gallica, 147(2): 165-172. Stévart T. & Geerinck D. 2000. Quatre nouveaux taxons d’Orchidaceae des genres Liparis, Bulbophyllum, Polystachya et Calanthe découverts à São Tomé et Príncipe. Syst. Geogr. Pl. 70: 141-148. Cribb P. J., la Croix I. & Stévart T. 1999. A new Polystachya (Orchidaceae) from São Tomé. Kew Bull. 54: 999-1001.