Wendy Applequist

Associate Curator

William L. Brown Center
Missouri Botanical Garden
P.O. Box 299
St. Louis, MO 63166-0299

office phone: + 1-314-577-0267;
fax: + 1-314-577-0800
email: wendy.applequist@mobot.org


1999- PhD (Botany), Iowa State University
1994- BS (Biology, cum laude), University of Illinois at Urbana-Champaign

Adjunct Instructor, Washington Univ. St. Louis

Curriculum vitae

General Research Interests

Medicinal Plants (authentication, quality control, taxonomy, chemical variability)
Natural Products Discovery
Taxonomy (Dombeya, Portulacaceae, Salicaceae s.l.; primarily in Madagascar)

Current Projects

Revisionary studies of Malagasy genera of Salicaceae s.l., especially Homalium
Management of natural products discovery programs
Participation in MU Botanical Center research, primarily on elderberry
Service as Secretary of the Nomenclature Committee for Vascular Plants

Publications (2005-2014)


Applequist, W. L. In press. A taxonomic revision of Dombeya section Decastemon (Malvaceae). Annals of the Missouri Botanical Garden.

Applequist, W. L. In press. Conioselinum. Flora of North America.

Applequist, W. L. In press. Ligusticum. Flora of North America.

Applequist, W. L. In press. A brief review of recent controversies in the taxonomy and nomenclature of Sambucus nigra sensu lato. Acta Horticulturae.

Applequist, W. L. In press. Legal and ethical issues related to DNA banking. In W. L. Applequist and L. M. Campbell, eds., DNA banking for the 21st century: Proceedings of the U.S. Workshop on DNA Banking. William L. Brown Center, St. Louis, MO.

Applequist, W. L. In press. Using traditional taxonomy and vouchers in authentication and quality control. In K. Reynertson and K. Mahmood, eds., Botanicals: Methods and technologies for quality control. CRC Press, Taylor & Francis Group, LLC, Boca Raton, FL.

Applequist, W. L., and C. D. Bell. In press. Symphoricarpos. Flora of North America.

Applequist, W. L., P. B. Phillipson, and G. E. Schatz. In press. A synoptic revision of the Malagasy endemic genus Calantica (Salicaceae). Adansonia.

Blasiak, L. C., A. W. Schmidt, H. Andriamiarinoro, T. Mulaw, R. Rasolomampianina, W. L. Applequist, C. Birkinshaw, F. Rejo-Fienena, P. P. Lowry II, T. M. Schmidt, and R. T. Hill. 2014. Bacterial communities in Malagasy soils with differing levels of disturbance affecting botanical diversity. PLoS One 9(1): e85097. doi:10.1371/journal.pone.0085097

Applequist, W. L. 2013. A nomenclator for Homalium (Salicaceae). Skvortsovia 1:12–74.

Applequist, W. L. 2013. Report of the Nomenclature Committee for Vascular Plants: 65. Taxon 62: 1315–1326.

Applequist, W. L., and J. S. Miller. 2013. Selection and authentication of botanical materials for the development of analytical methods. Analytical and Bioanalytical Chemistry 405: 4419–4428.

Dai, Y., L. Harinantenaina, P. J. Brodie, C. Birkinshaw, R. Randrianaivo, W. Applequist, M. Ratsimbason, V. E. Rasamison, Y. Shen, K. TenDyke, and D. G. I. Kingston. 2013. Two antiproliferative triterpene saponins from Nematostylis anthophylla from the Madagascar dry forest. Chemistry & Biodiversity 10: 233–240.

Liu, Y., L. Harinantenaina, P. J. Brodie, J. D. Bowman, M. B. Cassera, C. Slebodnick, M. W. Callmander, R. Randrianaivo, E. Rakotobe, V. E. Rasamison, W. Applequist, C. Birkinshaw, G. P. Lewis, and D. G. I. Kingston. 2013. Bioactive compounds from Stuhlmannia moavi from the Madagascar dry forest. Bioorganic & Medicinal Chemistry 21: 7591–7594.

Applequist, W. L. 2012. Report of the Nomenclature Committee for Vascular Plants: 64. Taxon 61: 1108–1117.

Walker, K. M., and W. L. Applequist. 2012. Adulteration of selected unprocessed botanicals in the U.S. retail herbal trade. Economic Botany 66: 321–327.

Applequist, W. L. 2011. A revision of the Dombeya rigida complex (Malvaceae). Adansonia, sér. 3, 33: 249–262.

Applequist, W. L. 2011. Proposal to conserve the name Ligusticum californicum (Apiaceae) with a conserved type. Taxon 59: 1288–1289.

Applequist, W. L., and D. E. Moerman. 2011. Yarrow (Achillea millefolium L.): a neglected panacea? A review of ethnobotany, bioactivity, and biomedical research. Economic Botany 65: 209–225.

Thomas, A. L., W. L. Applequist, G. E. Rottinghaus, and J. S. Miller. 2011 [publ. 2012]. Black cohosh rhizome and phytochemical production in response to shading, spacing, and age. Acta Horticulturae (ISHS) 925: 175–183.

Applequist, W. L., M. Callmander, G. Davidse, A. Sennikov, M. Thulin, P. Vorster, and G. Yatskievych. 2010. Apportionment of institutional votes for the Nomenclature Section: a rebuttal to Smith & al. Taxon 59: 1567–1570.

Applequist, W. L. 2009. A revision of the Malagasy endemic Helmiopsis (Malvaceae s.l.). Annals of the Missouri Botanical Garden 96: 521–540.

Applequist, W. L. 2009. Two new species of Dombeya (Malvaceae) from Madagascar. Novon 19: 289–294.

Applequist, W. L., H. Johnson, and G. Rottinghaus. 2008. (+)-Catechin, (-)-epicatechin, and gallic acid content of seeds of hybrid grapes hardy in Missouri. American Journal of Enology and Viticulture 59: 98–102.

Thomas, A. L., P. L. Byers, C. E. Finn, Y.-C. Chen, G. E. Rottinghaus, A. M. Malone, and W. L. Applequist. 2008. Occurrence of rutin and chlorogenic acid in elderberry leaf, flower, and stem in response to genotype, environment, and season. Acta Horticulturae (ISHS) 765: 197–206.

Applequist, W. L., D. J. McGlinn, M. Miller, Q. G. Long, and J. S. Miller. 2007. How well do herbarium data predict the location of present populations? A test using Echinacea species in Missouri. Biodiversity and Conservation 16: 1397–1407.

Hildreth, J., E. Hrabeta-Robinson, W. Applequist, and J. Betz. 2007. Standard operating procedure for the collection and preparation of voucher plant specimens for use in the nutraceutical industry. Analytical and Bioanalytical Chemistry 389: 13–17.

Thomas, A. L., G. E. Rottinghaus, J. K. Tracy, W. L. Applequist, B. E. Schweitzer, L. J. Havermann, S. F. Woodbury, J. S. Miller, M. R. Ellersieck, R. J. Crawford, Jr., and D. Gray. 2007. Occurrence of 23-epi-26-deoxyactein and cimiracemoside A in various black cohosh tissues throughout the growing season. HortScience 42: 535–539.

Applequist, W. L. 2006. Proposal to reject the names Plantago psyllium and Plantago cynops (Plantaginaceae). Taxon 55: 235–236.

Applequist, W. L., B. Avula, B. T. Schaneberg, Y.-H. Wang, and I. A. Khan. 2006. Comparative fatty acid content of seeds of four Cucurbita species grown in a common (shared) garden. Journal of Food Composition and Analysis 19: 606–611.

Applequist, W. L., and V. Lebot. 2006. Validation of Piper methysticum var. wichmannii (Piperaceae). Novon 16: 3–4.

Applequist, W. L., W. L. Wagner, E. A. Zimmer, and M. Nepokroeff. 2006. Molecular evidence resolving the systematic position of Hectorella (Portulacaceae). Systematic Botany 31: 310–319.

Miller, J. S., and W. L. Applequist. 2006. Research grade botanicals: issues in dietary supplement research. Acta Horticulturae 720: 49–58.

Thomas, A. L., R. J. Crawford, Jr., L. J. Havermann, W. L. Applequist, B. E. Schweitzer, S. F. Woodbury, and J. S. Miller. 2006. Effect of planting depth, planting season, and fungicide treatment on establishment of black cohosh in a poorly drained soil. HortScience 41: 374–376.

Applequist, W. L. 2005. A revision of the Malagasy endemic Talinella (Portulacaceae). Adansonia, sér. 3, 27: 47–80.

Applequist, W. L. 2005. The identity and typification of Smilax grandifolia Regel and S. regelii Killip & Morton. Taxon 54: 144–146.

Applequist, W. L. 2005. Root anatomy of Ligusticum species (Apiaceae) sold as osha compared to that of potential contaminants. Journal of Herbs, Spices and Medicinal Plants 11(3): 1–11.

Applequist, W. L., and D. B. Pratt. 2005. The Malagasy endemic Dendroportulaca (Portulacaceae) is referable to Deeringia (Amaranthaceae): molecular and morphological evidence. Taxon 54: 681–687.


Applequist, W. L., and L. M. Campbell, editors. In press. DNA banking for the 21st century: Proceedings of the U.S. Workshop on DNA Banking. William L. Brown Center, St. Louis, MO.

Applequist, W. 2006. The identification of medicinal plants: a handbook of the morphology of botanicals in commerce. American Botanical Council, Austin, TX.


Rainer Willi Bussmann
Jan Salick
Armand Randrianasolo
Wendy Applequist
Robbie Hart

Research Specialists
Tefy H. Andriamihajarivo
Miandry Fagnarena
Ashley Glenn
Jessie Griffard
Fortunat Rakotoarivony
Nivo H. Rakotoarivelo
Tabita Randrianarivony
Lucien Rasoaviety
Aina Razanatsima
Andrew Townesmith

Research Assistants
Taryn Pelch

Research Associates
Trish Flaster
Sarah K. Khan
Will McClatchey
Betty Millan Salazar
Narel Paniangua
Douglas Sharon
Carolina Tellez
Will Tuladhar-Douglas
Ina Vandebroek
Carlos Vega
Karen Meyer Walker